Relations between nature and capital have been a longstanding concern in the social sciences. Going beyond antinomies of posthumanist and political economic enquiry, this paper advances a set of relational analytics for incorporating liveliness into critical analyses of capital. Firstly, developing the concept of animal work, it shows how metabolic, ecological and affective labour become a productive economic force. Secondly, animating the commodity, it demonstrates how lively forces influence commodification and exchange, enabling or hindering accumulation. Thirdly, tracking animal circulation, it examines the logics of rendition that transform nonhuman life into capital. In conclusion, the paper develops a relational grammar for anatomizing the nature-capital dynamic, one that reorients the economic to be co-constituted by the ecological from the outset.