This paper seeks to rethink the urban as an ecological formation. It argues that contrary to an emphasis on the built environment, articulated through refrains of capital, planning and design, cities are lived achievements, emerging through fabrications between human and other-than-human forces. Ecological formations are fleshed out in three modes: ecologies that involve the cultivated, feral and the wild. There are distinct forms of urban governance affiliated with each, taken as hybrids of biopolitical and vernacular practices. Rethinking cities as ecological formations enables a more radical understanding of the difference and heterogeneity of urban worlds than those on offer in urban theory. It signals a shift from animating urban geographies towards understandings of how other-than-human geographies are constitutive of urban worlds.