#NEXUS [please cite Asher RJ, Maree S, Bronner G, Bennett NC, Bloomer P, Czechowski P, Meyer M, Hofreiter M. In Press. A phylogenetic estimate for golden moles (Mammalia, Afrotheria, Chrysochloridae). BMC Evolutionary Biology. For graphic images of character states, see project P299 on www.morphobank.org, "matrix" tab, "view matrix", "chars" button] BEGIN TAXA; TITLE Untitled_Taxa_Block; DIMENSIONS NTAX=30; TAXLABELS Elephantulus Echinops Geogale Hemicentetes Limnogale Microgale Micropotamogale Oryzorictes Potamogale Procavia Setifer Tenrec Ambly_corriae Ambly_hott Ambly_marleyi Ambly_robustus Ambly_septentrion Calco_obtusi Chlorot_leuco Chlorot_arendsi Chlorot_duthiae Chlorot_sclateri Chryso_asiatica Chryso_stuhlmanni Cspalax_trevel Cspalax_villosus Crypto_wintoni Eremi_granti Neambly_gunn Neambly_julian; END; BEGIN CHARACTERS; TITLE Untitled_Character_Matrix; DIMENSIONS NCHAR=144; FORMAT SYMBOLS= " 0 1 2 3 4 5 6 7 8" MISSING=? GAP=- ; CHARSTATELABELS 1 tymp_proc_basisphen_AH21 / 0_flat 1_contrib_bulla, 2 piriform_fenestr_AH22 / '0_abs-solid' '1_pres-open', 3 CTPP_AH23 / 0_small 1_contrib_post_bulla, 4 RTPP_AH24 / 0_pres 1_reduced, 5 epitymp_recess_AH26 / 0_contin_midEar 1_enlarged, 6 entotympanic / 0_reduced 1_shields_tubal_canal 2_contrib_bulla, 7 entotymp_shape / 0_lat_elongate 1_lat_pointed 2_lat_rounded, 8 tubal_canal_AH27 / 0_open 1_closed, 9 ectotymp_exposure_AH28 / 0_apheneric 1_semiphaneric 2_phaneric, 10 ectotymp_expansion_AH29 / 0_ring 1_expanded, 11 ectotymp_eartube / 0_short 1_vent_midpt_glenoid, 12 malleus_af18 / 0_small 1_enlarged '2_globular-large' 3_club, 13 temporal_bulla / 0_pres 1_abs, 14 inf_stap_foramen_AH31 / 0_abs 1_pres, 15 inf_stapedial_fmn / 0_separate_fmOvale 1_adjacent_fmOvale 2_confluent_fmOvale, 16 for_ovale / 0_separate_sphenfis 1_adjacent_sphenfis, 17 carotid_fmn / 0_antlat_jugfm 1_ant_jugfm, 18 ascpharyngeal_fmn / 0_posterior 1_medial_bulla, 19 craniopharyng_fmn / 0_abs 1_pres, 20 jugFm / 0_exposed_ventral 1_dorsal_occip_cond, 21 basisphen_pit_AH34 / 0_flat 1_pres, 22 fenestrate_basioccip_AH35 / 0_solid 1_open, 23 'mastoid-tubercle PM AH37' / 0_abs 1_petromast_pres, 24 'mastoid tubercle-squam AH38' / 0_abs '1_pres-squam', 25 squam_length_basicr_AH39 / 0_short 1_long, 26 entoglenoid_process_pres_AH50 / 0_abs 1_pres, 27 entoglen_jaw_artic_AH51 / 0_reduced 1_pres, 28 entoglen_composition_AH52 / 0_squamosal '1_squam-alisphen' 2_alisphen, 29 ethmoid_foramen_AH46 / 0_ant_sphenorb_fissure 1_within_sphenorb_fissure, 30 sphenorbFis_orientation / 0_opens_anteriorly 1_opens_laterally, 31 alisph_canal_pres_AH32 / 0_abs 1_pres, 32 alisph_canal_position_AH33 / 0_lat_sphenorb_fissure 1_contin_sphenorb_fis, 33 optic_foramen_AH44 / 0_pres 1_abs, 34 optic_foramen_size_AH45 / 0_similar_V 1_smaller_V, 35 suboptic_foramen_AH36 / 0_abs 1_pres, 36 maxilla_in_orbit_AH55 / 0_reduced 1_pres_no_frontal_contact 2_pres_frontal_contact, 37 external_petromast_AH43 / 0_post_nuchal_crest 1_ant_nuchal_crest 2_occluded, 38 lambd_plate_ap23 / 0_abs 1_temp_scar_anterior, 39 sagittal_crest_ap26 / 0_abs 1_pres, 40 occiput / 0_90angle 1_mushroom, 41 sinus_canal_AH47 / 0_pres 1_abs, 42 sinus_canal_foramen_AH48 / 0_confl_sphenorb_fissure '1_betw_sphenorb-ethmoid' 2_confl_ethmoid, 43 frontal_proportions_AH61 / 0_elongate 1_similar_parietal 2_shorter_parietal, 44 crista_galli_AH65 / 0_pres 1_abs, 45 dual_ectopterygoid_AH40 / 0_single_pair 1_pair_on_each_side, 46 maxillary_proc_pterygoid_AH41 / 0_reduced 1_past_posterior_toothrow, 47 'sphenopal-dors palatine fmna AH49' / 0_common_recess 1_separate, 48 postpal_spine_AH42 / 0_small 1_large, 49 palatal_length / '0_>_interEAM' '1_<=_interEAM', 50 gr_palatine_boss / 0_flat 1_defines_grpal_medial, 51 zygomatic_arch_AH56 / 0_present 1_reduced, 52 zyg_arch_size_AH57 / 0_robust 1_slender, 53 max_zygoma_origin_AH58 / 0_M1 1_M2 2_M3, 54 max_zygoma_root_size_AH59 / 0_large 1_reduced, 55 lacrimal_foramen_AH53 / 0_abs 1_pres, 56 lacrimal_foramen_position_AH54 / 0_in_orbit_ 1_anterior_orbit, 57 lacrimal_flange_AH64 / 0_smooth 1_rugose 2_projection, 58 infraorb_canal_length_AH60 / 0_longer_than_1_cheek_tooth '1_similar-smaller_1_cheek_tooth', 59 post_rostrum / 0_medial_ant_zygarch 1_lateral_ant_zygarch, 60 nasal_shape_AH62 / 0_broad_posteriorly 1_tapered_posteriorly, 61 nasal_length_AH63 / 'post_edge_at-anterior_orbit' 1_post_edge_posterior_antorbit, 62 antlat_process_pmax / 0_abs 1_pres, 63 size_upper_dentition_AH66 / '0_anterior-most_enlarged' 1_similar, 64 premax_dentition_AH67 / 0_zero 1_one 2_two 3_three 4_five, 65 inter_I1_gap_AH126 / 0_narrow 1_wide, 66 upper_antemolars_AH68 / 0_five 1_four 2_three 3_two, 67 upper_canine_AH69 / 0_similar_antemolars 1_bulbous 2_trenchant, 68 C_metastyle / 0_reduced 1_postbuc_projects, 69 P2_protocone / 0_pres 1_abs, 70 P3_protocone_af105 / 0_pres 1_abs, 71 P3_parastyle / 0_flush_ant_molar_wall 1_ant_projects 2_projects_P3_longer_than_BLwide, 72 P3_shape_AH70 / 0_narrow 1_molariform, 73 P4_shape_AH71 / 0_narrow 1_molariform, 74 P4_carnassial_AH72 / 0_no_shearing 1_shearing, 75 metaconeAH73 / 0_pres 1_connate 2_reduced, 76 protocone_AH74 / 0_pres 1_reduced, 77 hypocone_AH75 / 0_pres 1_reduced, 78 M1_parastyle_AH76 / 0_flush_ant_wall 1_anterior_projection, 79 centrobuccal_cleft_AH77 / 0_flat 1_pres, 80 M3_pres_AH78 / 0_pres 1_abs, 81 M3_stylar_shelf_AH79 / 0_narrow 1_broad, 82 M3_mesostyle_AH80 / '0_lingual-adjacent_paracone' 1_buccal, 83 M3_shape / 0_as_M2 1_half_M2, 84 lower_ant_dentition_AH81 / 0_larger_adjacent_teeth 1_similar_adjacent_teeth 2_smaller_adjacent_teeth, 85 lower_antemolars_AH82 / 0_eight 1_seven 2_six 3_five 4_three, 86 p3_shape_AH83 / 0_narrow 1_molariform, 87 p4_shape_AH84 / 0_narrow 1_molariform, 88 p4_talonid_AH85 / 0_pres 1_abs, 89 molar_talonids / 0_pres 1_abs, 90 hypoconulid_AH86 / 0_abs 1_pres, 91 'hypoflexid-paracone AH87' / 0_no_occlusion 1_occlusion, 92 secondary_hyoflexid_basin_AH88 / 0_pres 1_abs, 93 m3_pres_AH89 / 0_pres 1_abs, 94 m3_length_AH90 / 0_longer_antemolars 1_similar_antemolars 2_shorter_antemolars, 95 mand_condyle_length_AH92 / 0_globular 1_transverse, 96 mandib_condyle_number_AH93 / 0_single_artic 1_dual_artic, 97 internal_coronoid_fossa_AH95 / 0_smooth 1_pocketed, 98 mand_coronoid / 0_flat 1_dors_projects, 99 angular_process_ap91_af116 / 0_narrows_distally 1_broadens_distally, 100 axis_keel_AH96 / 0_pres 1_ventrally_flat, 101 sacral_verts_AH97 / '0_two-four' '1_>five', 102 sacroiliac_artic_AH98 / 0_S1 '1_S1-2' '2_S2-3', 103 sacral_neural_spines_AH99 / 0_no_fusion 1_fused_longitudinally, 104 clavicle_AH100 / 0_pres 1_short 2_abs, 105 first_rib / 0_narrow_riblike '1_robust-broad', 106 sternal_keel / 0_flat '1_ant-inf_projects', 107 scapular_shape_AH101 / 0_triangular 1_elongate, 108 scap_metacromion_pres_AH102 / 0_pres 1_abs, 109 scap_metacrom_shape / 0_blunt 1_elongate, 110 scap_acromion_pres_AH104 / 0_pres 1_abs, 111 scap_acromion_shape_AH105 / 0_blunt 1_elongate, 112 scapular_spine_length_AH106 / 0_lat_glenoid 1_at_glenoid, 113 root_scap_spine / 0_open 1_acute_angle, 114 hum_head / 0_sphere 1_DV_elongate, 115 deltoid_trough / 0_medial_convex '1_trough-medial_concave', 116 med_epicondyle_pres_AH108 / 0_reduced 1_pres, 117 dist_hum_breadth / 0_medEpiCond_small '1_50-75%_hum_len' '2_>85%_hum_len', 118 entepicond_fm_humerus_AH107 / 0_pres 1_abs, 119 humerus_olecranon_fenestr_AH110 / 0_ossified '1_fenest-thin', 120 distal_ulna_AH111 / 0_carpal_artic 1_no_carpal_artic, 121 ulnocarp_artic_AH112 / 0_broad 1_styloid_only, 122 forarm_longbones_AH113 / 0_two 1_three, 123 digital_pad_1&2 / 0_abs 1_pres, 124 manual_digit_1 / 0_vestigial 1_small 2_metac_over_half_MC3 3_hypertrophied, 125 manual_digit_2 / 0_vestigial 1_small 2_metac_over_half_MC3 3_hypertrophied, 126 manual_digit_3 / 0_vestigial 1_small 2_metac_over_half_MC2 3_hypertrophied, 127 manual_digit_4 / 0_vestigial 1_small 2_metac_over_half_MC3 3_hypertrophied, 128 manual_digit_5 / 0_vestigial 1_small 2_metac_over_half_MC3 3_hypertrophied, 129 'number rib-bearing verts' / 0__13 1_14 2_15 3_16 4_17 5_18 6_19 7_20 8_21, 130 caudal_verts / 0_centra_similar_presacrals 1_centra_smaller_presacrals, 131 iliac_blades_AH114 / '0_round-sagittal' 1_coronal, 132 ilia_cranial_extent_AH115 / 0_past_sacroiliac_artic 1_even_sacroiliac_artic, 133 iliopectineal_tubercle_AH116 / 0_smooth 1_pres, 134 iliopect_tubercle_shape_AH117 / 0_medial 1_ventral, 135 internal_pelvic_shape_AH118 / 0_flat 1_concave, 136 pelvic_obt_fmn / 0_larger_acetab 1_smaller_acetab 2_abs, 137 pubic_symphysis_AH119 / 0_broad 1_narrow 2_separate, 138 pubis_angle / '0_>15_from_ilium' 1_parallel_ilium, 139 pubis_length_AH120 / 0_similar_ischium 1_longer_ischium, 140 fem_3rd_trochanter_AH122 / 0_small_proximal 1_reaches_midpoint_shaft, 141 prox_fibula / 0_narrow 1_AP_process, 142 'tibia-fibula AH123' / 0_unfused 1_fused, 143 distal_fibula_AH124 / 0_flat 1_concave '2_ball-condyle', 144 calcaneal_tubercle_AH125 / 0_blunt 1_pointed ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 ] [ . . . . . . . . . . . . . . ] Elephantulus 00100221211011001-100000010100(01)0001010000111101100001010110100130000000110000001---10010000-(01)200010101000?0010010000-011-00122220000111000000100 Echinops 100100-020-011001-1010010111100-0102100000010100001-0011010110020120111110211101---2211100111-000101010000000001000100(01)010022222?1010-0010000001 Geogale 111000-020-011001-000(01)01011000110112100000110100001-111100011002110011-01021111010003011001100000100010000000001000101101002222240000-001010000? Hemicentetes 1(01)0100-020-010-01-100001011110110102000000(01)10100001-20110101001301(02)011100021111011011001000-01000101010000000001000100001001222231110-0010100001 Limnogale 111000-020-01(01)001??00100011000110112100001210100001-100-0101100301001011102111101002101100110010010001000?00000100010000100?222??0000-0010100101 Microgale 111000-020-011001-10000001100011011210000(01)110100001-10110101100301(02)010101021111010021011001100100101010000000011000100001002222230000-0010100101 Micropotamogale 101000-110-010-011100101011000110112100001210100001-100-01011003010010111020111010021(01)1100110110010001020?01-1-1?0?1010010012222?000110011100101 Oryzorictes 111000-020-011001-000000011000110102100000110100001-101(01)010100130120111010211110100110110011001001010100000000010001000010012222(345)0000-0010100101 Potamogale 101000-110-010-0111001010110001101(01)010001-2(01)0100001-100-0101000301001011101011101002101000100(01)10010001020001-1-10000-10000022222(34)000110010110101 Procavia 01010220211010-01-00000011110011000020001-101(01)01000000102000000101010011100000000-022110000-011001-011120001-1-10000-1100000222281100-0000000000 Setifer 100100-020-011001-101001011110110102100000010100001-(01)01101011002(01)12011111021100011112111001102000101010000000001000100(01)010022222(34)1010-0010100001 Tenrec 1(01)0(01)00-020-010-011101001011110110102001000110100001-10110101001(23)01201111102110001101(12)11100110(01)0001010100000000010001001010012222(67)1110-0010100001 Ambly_corriae 101?11102100111000(01)10000010-100-1-0?0001????00000001001(01)010??1030100101110201011---2111000111-00001??????????????????000?10123006??????????????? Ambly_hott 101011102100111000010000010-100-1-0?0001???1000000010010010??1030100102110201011---2111000111-000011011011001000011120000101230061000-0121111121 Ambly_marleyi 101??110210?111000110000010-100-1-0?0001????000000010010010111030100112110201111---2111000111-00001??????????????????000?1??????6??????????????? Ambly_robustus 101?1110210?111000110000010-100-1-0?0001????00000001001(01)010??1030100101110201011---2111000111-00001??????????????????000?10123006??????????????? Ambly_septentrion 101??110211?111000010000010-100-1-0?0001????000000010010010??1030100101110201011---2111000111-0000110110110010000111200001?1230061000-01211?1121 Calco_obtusi 101011102100111000010000010-100-1-0?0001???1000000010011010??1030101110110201001-1-2111110111-000111021011001000011120000101230061000-01211111?? Chlorot_leuco 101011102101112000010000010-100-1-020001???100000001001101010103010011011021100011121111101102000111021011001000011110000101230061000-012111112? Chlorot_arendsi 101011102101111000110000010-100-1-0?0001???1000000010011010??1030100100110201000111211100011020001110110110010000111200001012300?1000-012111112? Chlorot_duthiae 101011102101111100010000010-100-1-0?0001???1000000010010010??1030100101110201010101211100011020001110110110010001111200001012300?1000-0121111121 Chlorot_sclateri 101011102101111000010000010-100-1-0?0001???1000000010010010??103010010111020101(01)10121110001102000111011011001000111120000101230061000-0121111121 Chryso_asiatica 101011002103012111010000010-100-1-0?0001???10000000100100101?1030100101110201000101211111011020001110110110010000111200001012300610(01)0-01211111?1 Chryso_stuhlmanni 101011002103012111010000010-100-1-0?0001???100000001001?010??10301001001102010?01-12111?00110200011?0?10110010000111200001012300610?0-0121?111?? Cspalax_trevel 10101120210201211(01)11(01)000010-100-1-020111???1000(01)01010011010??103010010111020101011121110001102000111011011000000011110000101230071000-0121011121 Cspalax_villosus 101011202102012110110000010-10(01)11-0201(01)1???1000000010011010??10301001001102010101012111000110200011?0?10110000000111100001012300?10?0-0121?111?? Crypto_wintoni 101011102103012100010000010-100-1-0?0001???1000000010011011??103010010(01)1102010101012111110110200011?0?10??0???00???12000011223106?0?0-0?2??1?1?? Eremi_granti 101011102102012100000000010-010-1-0?0001???100001001001?010??103010011(01)1102010101012111110110200011?0?10110010001101100001022210410?0-0121?111?? Neambly_gunn 101011102100111000010000010-100-1-0?0001???1000000010011010??103010010111020101(01)0-121110101102000011011011001000011120000101230061000-0121111121 Neambly_julian 101011102100111000010000010-100-1-0?0001???1000010010011010??10301001011102010(01)1---2111110?1(01)-000011011011001000011120000101230071000-0121111121 ; END; BEGIN paup; pset mstaxa=polymorph collapse=minbrlen; END;