Acknowledgements: This work is part of a project that has received funding from the European Union's Horizon 2020 research and innovation programme under the Marie Skłodowska-Curie grant agreement No. 813786 (EVOdrops). This work reflects only the author's view, and the Research Executive Agency is not responsible for any use that may be made of the information it contains. It was supported by a studentship from the BBSRC Doctoral Training Account, (E. J. M., BB/M011194/1), a Trinity College/Benn W Levy SBS DTP PhD Studentship (M. G.) and an H2020 ERC Advanced Investigator Award (F. H., 695669).